I was watching a squirrel happily burying nuts in the garden and I have always been amazed at how they know where they buried them. Now I'm pretty sure that this squirrel knows exactly where they are as when it snows you can see the tracks directly to a particular point and not tracks all over the place as it would be if he was searching. Anyway someone claimed that they don't know where they are they do lots of searching to find them again. Well I knew from watching my squirrel, that comment wasn't true. So I did some searching......
According to the study "Grey Squirrels Remember the Locations of Buried Nuts," published in Princeton University's journal "Animal Behavior," squirrels use spatial memory often to locate stored food. The study shows squirrels go back more often to their own food caches than to the caches of other squirrels, suggesting that they use memory to locate their food. Squirrels often bury their food near landmarks that aid them in remembering where they stored the food. The study also suggests that squirrels bury food in a series of locations that help them form a cognitive map of all storage locations.
A seminal 1991 research paper published in the journal Animal Behavior showed that even when multiple grey squirrels (Sciurus carolinensis) bury their stash in close proximity to one another, individuals of this species will remember and return to the precise locations of their personal cache. This is echoed by multiple other studies, showing that the squirrels' spatial memory helps them map out the territory around them to find their food. Under certain conditions — like when their nuts are buried under snow — a sense of smell won’t always be effective in helping them find food. So, it makes sense that squirrels could be relying on other cues.
LINKS
https://animals.mom.me/squirrels-really-bury-food-11108.html
https://www.livescience.com/64104-how-do-squirrels-find-buried-nuts.html
https://www.wildlifeonline.me.uk/questions/answer/what-controls-the-caching-behaviour-of-squirrels-and-how-do-they-find-their-bur